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For anyone that finds this useful, this is my take on the two, after owning 3 previous 5th gens and now a 6th gen (for the last month with 2,000 miles on it including towing miles). Additionally, I have owned 1st and 2nd gen Tundras, a 200 series, a GX 470 and 460, a 1st and 2nd gen 4Runner... I like Toyotas.

5th Gen:

The 5th Gen 4Runner is an absolute beast of a vehicle. The 1GR-FE is as bulletproof as Toyota engines get. Likewise the 20 year old A750 5 speed almost never failed. The word 'bulletproof' is the embodiment of the 5th gen 4Runner. As a former owner of the platform, there were only two downsides to me (a 40 year old, 'overlander' that cares more and more about safety after being in a serious auto accident last year and not about power and going fast):

1. The 5th gen was very picky about tire balancing and weight. All three 5th gens I owned, and thousands on all the forums, had issues when bigger tires were mounted, and suffered from wheel shake. This included 2 brand new units I had with less than 10 miles on them when switching to bigger all terrains (even modest 265/70/R17). This has been completely fixed on the 6th gen.

2. Crash safety: older vehicles simply don't perform as well and the 5th gen (same with the GX 460) left a lot to be desired in moderate offset and front end collisions, leaving the driver footwell very compromised: https://www.iihs.org/ratings/vehicle/toyota/4runner-4-door-suv/2022. The 6th gen, given that it is structurally similar if not identical to the 2024 tacoma all the way to the cab, has improved crash testing significantly. You can have a lot of problems in life, until you have a health problem, then you only have one problem. Safety is crucial, and I'm glad to see Toyota engineers improved the platform in crash safety.

That's it, I can find no other complaints on the 5th gen, I loved all of mine, and probably would still own one if it weren't for the vibrations that I was getting through the steering wheel every time I tried to mount larger all-terrain tires.

6th Gen:

The 6th Gen is a stark departure from the 5th gen. It is very close, if not identical to the Tacoma in every mechanical way (the 4Runner is thankfully still made in the Tahara factory in Japan, likely the best automotive plant on the planet). It features a 4cylinder turbo with hybrid option. It is no longer only port injected, but now includes D4S, Toyota's port and direct injection system, that in other models, works very well and is very proven.

As someone that very much liked the 5th gen drivetrain and did not think it was underpowered (despite living at 5,300 feet above sea level) for a midsize SUV, I can tell you that the 6th gen drivetrain (in terms of drivability, not necessarily durability and longevity) is a vast improvement. And again, I love the 4.0 V6. The 2.4 Turbo (mine is gas, SR5) and 8 speed are fantastic. There is occasionally a small hesitation/rough shift between gears 2 and 3, but it's not that annoying and only occurs to me under spirited acceleration.

On a recent drive from Boulder Colorado to Lander Wyoming (a route I drive regularly for work), and a route that previously in my 5th gen, was always filled with downshifting and 4,000 RPM, now is quiet and effortless. The 6th gen almost never even downshifts, and when it does, I stay at speed well under 3,000 RPM. The real world torque is very noticeable, and such a pleasant experience and the 8 speed gearbox is a welcomed addition. I also greatly appreciate the 6 grease zerks found on the front and rear drive shaft. I can't recall if these were present on my 5th gen, but either way, serviceability is a win in my book.

As soon as I got it, I added my favorite tire, the Falken Wildpeak AT4W, in a modest 265/70/R17 in SL variety (this is my commuter so no need for a C rated or E rated tire). Prior to doing this, the 4Runner was achieving (again at altitude, where I always experience better fuel economy than at lower altitudes) 26.1 mpg hand calculated (although the computer is almost identical and is very accurate in my experience). After the tire upgrade, I've been averaging around 22 mpg, so a significant hit, but still a very good number for a 4,600 pound SUV with all terrains (note that I try and drive very easy, this is also on 91 octane fuel). The air dam will come off soon:

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective IMG_3198


Towing with the new 4Runner is equally is enjoyable. The power and low speed thrust is considerable. I towed my 7x14 cargo trailer with a couch and some other furniture about 50 miles (trailer probably weighed close to 4,000 lbs) and it pulled very well; much, much better than my 5th gen.

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective IMG_3198



I appreciate the SR5 coming with 17 inch wheels and wish higher trims would do the same. 18s, 19s and 20s, do not belong on an SUV intended for offroad use in my opinion.

The user experience of the 6th gen has been delightful and I'm happy I made the switch. Many people complain about materials, but the plastic dashboard and armrests by the window do not bother me, and again, as a rugged all terrain machine, I prefer more durable materials that can take more of a beating.

The only annoyance so far on the 6th gen is that the plastic body moulding on the rear tires fills with mud, and even after a thorough car wash, it was still there. I had to clean it by hand while opening the door. But again, this is a very small thing.

Style is subjective, and I do prefer the 5th gen more than the 6th, but things have to evolve I guess.

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective IMG_3197 (1)


If you have any questions about the comparison, let me know. I think Toyota did well. Yes it has a 4 cylinder, and a turbo, but servicing this engine will be very easy (spark plugs should take less than 10 minutes, which is great, given that they are called for every 40,000 miles in the 6th gen), and the driving experience is far better than the outgoing model.
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spyder40

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Looking to swap my 2021 ORP for a 2025 ORP Hybrid. Pull a trailer with a SxS at 3500-4000lbs and the locking in 4th/running at 3000+ rpm gets old. Good to hear the new one pulls better than the 5th gen
 
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YotaOverland

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Looking to swap my 2021 ORP for a 2025 ORP Hybrid. Pull a trailer with a SxS at 3500-4000lbs and the locking in 4th/running at 3000+ rpm gets old. Good to hear the new one pulls better than the 5th gen
I’m looking to do the same thing, haven’t bought the trailer or side by side (sold the cargo trailer yesterday), let me know how it goes if you end up switching.
 

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One thing I am surprised you didn't mention was the serious brake dive on the Gen 5. My 2018 Pro was sketchy as hell in a panic stop, all the weight transferred forward, and the rear end started moving around. Once you got comfortable with it you were okay, but it caught a few passengers out and certainly wasn't pleasant.

My 2018 also used to lean way over in the corners more than my new rig. I actually liked that, in tight corners, especially on loose surfaces, you could rotate the car with that weight shift. The new version is much more planted.
 

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One thing I am surprised you didn't mention was the serious brake dive on the Gen 5. My 2018 Pro was sketchy as hell in a panic stop, all the weight transferred forward, and the rear end started moving around. Once you got comfortable with it you were okay, but it caught a few passengers out and certainly wasn't pleasant.

My 2018 also used to lean way over in the corners more than my new rig. I actually liked that, in tight corners, especially on loose surfaces, you could rotate the car with that weight shift. The new version is much more planted.
The brake dive was terrible in my 2018 and the 2021, drove me nuts. Put Icon UCA's and Fox 2.5's on the 2021 and it was gone. Lock up the brakes and almost no dive at all. One of the things I checked on when I test drove a 2025 and it doesn't have it. Also got rid of the body sway with some adjusting of the DSC's. Truck rides like a dream now, flat in the corners and plain planted on stuff like washboards and good whoopdee bumps. If it wasn't for the towing I wouldn't get rid of it.
 

time2playandy

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For anyone that finds this useful, this is my take on the two, after owning 3 previous 5th gens and now a 6th gen (for the last month with 2,000 miles on it including towing miles). Additionally, I have owned 1st and 2nd gen Tundras, a 200 series, a GX 470 and 460, a 1st and 2nd gen 4Runner... I like Toyotas.

5th Gen:

The 5th Gen 4Runner is an absolute beast of a vehicle. The 1GR-FE is as bulletproof as Toyota engines get. Likewise the 20 year old A750 5 speed almost never failed. The word 'bulletproof' is the embodiment of the 5th gen 4Runner. As a former owner of the platform, there were only two downsides to me (a 40 year old, 'overlander' that cares more and more about safety after being in a serious auto accident last year and not about power and going fast):

1. The 5th gen was very picky about tire balancing and weight. All three 5th gens I owned, and thousands on all the forums, had issues when bigger tires were mounted, and suffered from wheel shake. This included 2 brand new units I had with less than 10 miles on them when switching to bigger all terrains (even modest 265/70/R17). This has been completely fixed on the 6th gen.

2. Crash safety: older vehicles simply don't perform as well and the 5th gen (same with the GX 460) left a lot to be desired in moderate offset and front end collisions, leaving the driver footwell very compromised: https://www.iihs.org/ratings/vehicle/toyota/4runner-4-door-suv/2022. The 6th gen, given that it is structurally similar if not identical to the 2024 tacoma all the way to the cab, has improved crash testing significantly. You can have a lot of problems in life, until you have a health problem, then you only have one problem. Safety is crucial, and I'm glad to see Toyota engineers improved the platform in crash safety.

That's it, I can find no other complaints on the 5th gen, I loved all of mine, and probably would still own one if it weren't for the vibrations that I was getting through the steering wheel every time I tried to mount larger all-terrain tires.

6th Gen:

The 6th Gen is a stark departure from the 5th gen. It is very close, if not identical to the Tacoma in every mechanical way (the 4Runner is thankfully still made in the Tahara factory in Japan, likely the best automotive plant on the planet). It features a 4cylinder turbo with hybrid option. It is no longer only port injected, but now includes D4S, Toyota's port and direct injection system, that in other models, works very well and is very proven.

As someone that very much liked the 5th gen drivetrain and did not think it was underpowered (despite living at 5,300 feet above sea level) for a midsize SUV, I can tell you that the 6th gen drivetrain (in terms of drivability, not necessarily durability and longevity) is a vast improvement. And again, I love the 4.0 V6. The 2.4 Turbo (mine is gas, SR5) and 8 speed are fantastic. There is occasionally a small hesitation/rough shift between gears 2 and 3, but it's not that annoying and only occurs to me under spirited acceleration.

On a recent drive from Boulder Colorado to Lander Wyoming (a route I drive regularly for work), and a route that previously in my 5th gen, was always filled with downshifting and 4,000 RPM, now is quiet and effortless. The 6th gen almost never even downshifts, and when it does, I stay at speed well under 3,000 RPM. The real world torque is very noticeable, and such a pleasant experience and the 8 speed gearbox is a welcomed addition. I also greatly appreciate the 6 grease zerks found on the front and rear drive shaft. I can't recall if these were present on my 5th gen, but either way, serviceability is a win in my book.

As soon as I got it, I added my favorite tire, the Falken Wildpeak AT4W, in a modest 265/70/R17 in SL variety (this is my commuter so no need for a C rated or E rated tire). Prior to doing this, the 4Runner was achieving (again at altitude, where I always experience better fuel economy than at lower altitudes) 26.1 mpg hand calculated (although the computer is almost identical and is very accurate in my experience). After the tire upgrade, I've been averaging around 22 mpg, so a significant hit, but still a very good number for a 4,600 pound SUV with all terrains (note that I try and drive very easy, this is also on 91 octane fuel). The air dam will come off soon:

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective {filename}


Towing with the new 4Runner is equally is enjoyable. The power and low speed thrust is considerable. I towed my 7x14 cargo trailer with a couch and some other furniture about 50 miles (trailer probably weighed close to 4,000 lbs) and it pulled very well; much, much better than my 5th gen.

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective {filename}



I appreciate the SR5 coming with 17 inch wheels and wish higher trims would do the same. 18s, 19s and 20s, do not belong on an SUV intended for offroad use in my opinion.

The user experience of the 6th gen has been delightful and I'm happy I made the switch. Many people complain about materials, but the plastic dashboard and armrests by the window do not bother me, and again, as a rugged all terrain machine, I prefer more durable materials that can take more of a beating.

The only annoyance so far on the 6th gen is that the plastic body moulding on the rear tires fills with mud, and even after a thorough car wash, it was still there. I had to clean it by hand while opening the door. But again, this is a very small thing.

Style is subjective, and I do prefer the 5th gen more than the 6th, but things have to evolve I guess.

2025 Toyota 4runner 6th gen vs 5th gen: My Perspective {filename}


If you have any questions about the comparison, let me know. I think Toyota did well. Yes it has a 4 cylinder, and a turbo, but servicing this engine will be very easy (spark plugs should take less than 10 minutes, which is great, given that they are called for every 40,000 miles in the 6th gen), and the driving experience is far better than the outgoing model.
Great write-up! Re: towing, what psi are you running? And what trailer brake controller? Thanks!
 
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YotaOverland

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One thing I am surprised you didn't mention was the serious brake dive on the Gen 5. My 2018 Pro was sketchy as hell in a panic stop, all the weight transferred forward, and the rear end started moving around. Once you got comfortable with it you were okay, but it caught a few passengers out and certainly wasn't pleasant.

My 2018 also used to lean way over in the corners more than my new rig. I actually liked that, in tight corners, especially on loose surfaces, you could rotate the car with that weight shift. The new version is much more planted.
That is a great point, I completely forgot about that, as I was just writing this over dinner last night. Yes the new one is much more surefooted.
 
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Great write-up! Re: towing, what psi are you running? And what trailer brake controller? Thanks!
I was just moving a few pieces of furniture then selling the trailer (bought it when I had my Superduty), so I didn't install a trailer brake controller, likely will in the future though. Tires were at 38 psi.
 

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As a current 5th and 6th gen owner, I have to say I very much look forward to hopping in the 6th gen versus the 5th. Both drive totally different, and in a good way. The big one for me was the low end torque and ease of the steering of the 6th. The 5th is just classic, smooth truck feel. I will own both for years to come.
 

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One thing I am surprised you didn't mention was the serious brake dive on the Gen 5. My 2018 Pro was sketchy as hell in a panic stop, all the weight transferred forward, and the rear end started moving around. Once you got comfortable with it you were okay, but it caught a few passengers out and certainly wasn't pleasant.

My 2018 also used to lean way over in the corners more than my new rig. I actually liked that, in tight corners, especially on loose surfaces, you could rotate the car with that weight shift. The new version is much more planted.
Indeed, The 5th gen nose dive is terrible...
"Planted" is the exact word that came to my mind when I test drove a 6th gen.
 

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Great comparo between the two @YotaOverland , I had an 18 OR that I loved and similarly went with an SR5 this time. The 4.0 was more than adequate and I never needed more necessarily. With our 6th gen though, maybe it's not much quicker objectively, but the usable power and mid-range improvement in the is rather significant and we just hope it's durable. In fact, it easily stacks up well compared to our GX460, engine sound excluded.

I'm glad you mentioned the tire balance issue! I went through a lot of effort in getting our 5th gen right on Terra Grappler G2's, AT3's, and KO2's. It was always a chore, a lot of rebalancing, and a lot of nagging to ensure techs were using the flange plate on the RF balancer, rotating the tire on the rim, etc. Fingers crossed, hopefully the suspension geometry and the TNGA-F platform is less susceptible and sensitive to this. So far, I have Open Country AT3's mounted and no issues so far through 2000ish miles.

Also, seems to be a common observation but the platform control on the 6th gen is a huge improvement, which is somewhat silly to say given this is a 15 year newer vehicle. The body movement is very well controlled and roll is rather stable for a BoF truck, I was pleasantly surprised prior to buying mine on the test drive, even compared to our GX460 with KDSS.

Anyway, loved our 5th gen also....hoping the 6th gen continues to impress.
 
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Great comparo between the two @YotaOverland , I had an 18 OR that I loved and similarly went with an SR5 this time. The 4.0 was more than adequate and I never needed more necessarily. With our 6th gen though, maybe it's not much quicker objectively, but the usable power and mid-range improvement in the is rather significant and we just hope it's durable. In fact, it easily stacks up well compared to our GX460, engine sound excluded.

I'm glad you mentioned the tire balance issue! I went through a lot of effort in getting our 5th gen right on Terra Grappler G2's, AT3's, and KO2's. It was always a chore, a lot of rebalancing, and a lot of nagging to ensure techs were using the flange plate on the RF balancer, rotating the tire on the rim, etc. Fingers crossed, hopefully the suspension geometry and the TNGA-F platform is less susceptible and sensitive to this. So far, I have Open Country AT3's mounted and no issues so far through 2000ish miles.

Also, seems to be a common observation but the platform control on the 6th gen is a huge improvement, which is somewhat silly to say given this is a 15 year newer vehicle. The body movement is very well controlled and roll is rather stable for a BoF truck, I was pleasantly surprised prior to buying mine on the test drive, even compared to our GX460 with KDSS.

Anyway, loved our 5th gen also....hoping the 6th gen continues to impress.
Thanks @dc503 Yeah, I feel you with my 21 TRD OR, and my 2022 SR5, I just eventually gave up and went back to stock sized tires, I got so tired of going to discount tire and chasing different variables to try to get the wheel to stop shaking. I'm now at 2000 miles as well on my sixth gen with Falken Wildpeaks, that I could never get to balance right on the 5th gen, and everything is good so far.

Yes, I'm a bit embarrassed that I completely forgot about how much of an improvement the platform control is, it's night and day better than the 5th Gen, and you're right, I like how flat it drives even compared to my old GX 460. I had a 130 lbs Go fast roof tent on the GX with KDSS, which I imagine will do better than the 6th gen 4runner with a tent up top, but it will still be a significant improvement over the 5th gen.
 

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I feel the same, i had a 5th gen for years but at sea level the best mpg i could get was 17-18mpg as a stock truck. Now i see 24-26mpg in the 6 gen if i stay under 70-75 mph. The Bilsteins are a bit rough but i also appreciate their body control over the 5th gen
 
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How does the 6g compare to the GX460 in your opinion? I was looking at these hard for a while up until recently but am now just about sold on the 6g. Would love to hear your input with the GX in mind. TIA!
Good question, I actually like the drivetrain of the 4runner better. Turbos just make all the difference for low end torque. The GX is way more luxurious of course, and it will get 17-18 mpg vs the 4runner’s 21-25. The lexus is quieter but its also much older tech. I love them both and was in your position and almost went with the GX but went with the 4runner and am happy I did. Good luck in your decision!
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